Archaeology and ichnology at Gombore II-2, Melka Kunture, Ethiopia: everyday life of a mixed-age hominin group 700,000 years ago.

We report the occurrence at 0.7 million years (Ma) of an ichnological assemblage at Gombore II-2, which is one of several archaeological sites at Melka Kunture in the upper Awash Valley of Ethiopia, 2000 m asl. Adults and children potentially as young as 12 months old left tracks in a silty substrate on the shore of a body of water where ungulates, as well as other mammals and birds, congregated. Furthermore, the same layers contain a rich archaeological and palaeontological record, confirming that knapping was taking place in situ and that stone tools were used for butchering hippo carcasses at the site. The site gives direct information on hominin landscape use at 0.7 Ma and may provide fresh perspective on the childhood of our ancestors

Beaver exploitation, 400,000 years ago, testifies to prey choice diversity of Middle Pleistocene hominins

Data regarding the subsistence base of early hominins are heavily biased in favor of the animal component of their diets, in particular the remains of large mammals, which are generally much better preserved at archaeological sites than the bones of smaller animals, let alone the remains of plant food. Exploitation of smaller game is very rarely documented before the latest phases of the Pleistocene, which is often taken to imply narrow diets of archaic Homo and interpreted as a striking economic difference between Late Pleistocene and earlier hominins. We present new data that contradict this view of Middle Pleistocene Lower Palaeolithic hominins: cut mark evidence demonstrating systematic exploitation of beavers, identified in the large faunal assemblage from the c. 400,000 years old hominin site Bilzingsleben, in central Germany. In combination with a prime-age dominated mortality profile, this cut mark record shows that the rich beaver assemblage resulted from repetitive human hunting activities, with a focus on young adult individuals. The Bilzingsleben beaver exploitation evidence demonstrates a greater diversity of prey choice by Middle Pleistocene hominins than commonly acknowledged, and a much deeper history of broad-spectrum subsistence than commonly assumed, already visible in prey choices 400,000 years ago.Human Origin

Dancing with wolves at Schöningen 13II-4

The Schöningen 13II-4 site has produced a wealth of insight into the hunting and butchery activities of Middle Pleistocene hominins, highlighted by the famous Schöningen spears preserved with hundreds of cut-marked and broken horse bones. The bones of carnivores are rare at the site, but tooth pits, scores, and other markings that record their presence are abundant. Here we describe the carnivore remains from Schöningen 13II-4 and provide a detailed analysis of carnivore markings on different skeletal parts in the faunal assemblage and their spatial distribution. In studying carnivore activities at Schöningen, we aim to achieve a more comprehensive view of site taphonomy and, in turn, a better appreciation of the anthropogenic process that shaped the archaeological record. The placement and sequence of carnivore marks on the bones in relation to butchery marks indicates that carnivores scavenged from the remains of hominin kills. In the large horse bone assemblage, carnivore damage is more prevalent on limb bones of juveniles than adults. This pattern reveals that adult horse carcasses were fully butchered by hominins, but juvenile horse carcasses were abandoned earlier in the butchery process, leaving more consumable tissues that attracted scavenging carnivores. Tooth pits and scores on the Schöningen remains are very large and compare well with markings produced by wolves, especially those observed in a sample of modern wolf-gnawed bones we collected and analysed from Adler- und Wolfspark Kasteelburg. Clusters of carnivore-damaged bones appear around the periphery of dense concentrations of bones butchered by hominins, suggesting that wolves displaced some skeletal elements quickly after abandonment by hominins. Such a spatial pattern hints at the long-standing co-habitation of the Schöningen landscape by hominins and wolves during the Middle Pleistocen

Widespread evidence for elephant exploitation by Last Interglacial Neanderthals on the North European plain

Neanderthals  hunted  and  butchered  straight- tusked  elephants,  the  largest  terrestrial  mammals of the Pleistocene, in a lake landscape on the North European plain, 125,000 years ago, as recently shown by a study of the Last Interglacial elephant assemblage from Neumark- Nord (Germany). With evidence for a remarkable focus on adult males and on their extended utilization, the data from this location are thus far without parallel in the archaeological record. Given their relevance for our knowledge of the Neanderthal niche, we investigated whether the Neumark- Nord subsistence practices were more than a local phenomenon, possibly determined by local characteristics. Analyzing elephant remains from two other Last Interglacial archaeological sites on the North European plain, Gröbern and Taubach, we identified in both assemblages similar butchering patterns as at Neumark-Nord, demonstrating that extended elephant exploitation was a widespread Neanderthal practice during the (early part of the) Last Interglacial. The substantial efforts needed to process these animals, weighing up to 13 metric tons, and the large amounts of food generated suggest that Neanderthals either had ways of storing vast amounts of meat and fat and/or temporarily aggregated in larger groups than com-monly acknowledged. The data do not allow us to rule out one of the two explanations, and furthermore both factors, short- term larger group sizes as well as some form of food preservation, may have played a role. What the data do show is that exploitation of large straight-tusked elephants was a widespread and recurring phenomenon amongst Last  Interglacial Neanderthals on the North European plain.Human Origin

Hunting and processing of straight-tusked elephants 125.000 years ago: implications for Neanderthal behavior

Straight-tusked elephants (Palaeoloxodon antiquus) were the largest terrestrial mammals of the Pleistocene, present in Eurasian landscapes between 800,000 and 100,000 years ago. The occasional co-occurrence of their skeletal remains with stone tools has generated rich speculation about the nature of interactions between these elephants and Pleistocene humans: Did hominins scavenge on elephants that died a natural death or maybe even hunt some individuals? Our archaeozoological study of the largest P. antiquus assemblage known, excavated from 125,000-year-old lake deposits in Germany, shows that hunting of elephants weighing up to 13 metric tons was part of the cultural repertoire of Last Interglacial Neanderthals there, over >2000 years, many dozens of generations. The intensity and nutritional yields of these well-documented butchering activities, combined with previously reported data from this Neumark-Nord site complex, suggest that Neanderthals were less mobile and operated within social units substantially larger than commonly envisaged.Human Origin

A detailed analysis of the spatial distribution of Schöningen 13II-4 'Spear Horizon' faunal remains

The Middle Pleistocene Schöningen 13II-4 'Spear Horizon' (Germany) is a key site for the study of human evolution, most notably for the discovery of Paleolithic wooden weaponry and evidence for developed hunting strategies. On the other hand, the 'Spear Horizon' offers an excellent opportunity to approach hominin spatial behavior, thanks to the richness of the archeological assemblage, its exceptional preservation, and the vast expanse of the excavated surface. Analyzing how space was used is essential for understanding hominin behavior at this unique open-air site and, from a wider perspective, for approaching how humans adapted to interglacial environments. In this article, we present an exhaustive spatial study of the complete Schöningen 13II-4 'Spear Horizon' faunal assemblage and its archeological context, combining zooarcheology and spatial analysis through the extensive application of geographic information systems. Our results indicate the existence of different activity areas related to changes in the position of the shoreline due to fluctuations of water table levels of the Schöningen paleolake. These activity areas were likely used on a seasonal basis, whereas the spatial patterning observed in the distribution of faunal remains suggests a diversity of behavioral strategies in terms of intensity and/or duration of occupations. This study refines previous interpretations of the site and reconstructs human behavioral adaptations and the occupational changing lakeland environment during the Middle Pleistocene in Europe.This research is part of the ‘Hominin subsistence strategies during the Middle Pleistocene in Schoningen € ’ project, funded by the Deutsche Forschungsgemeinschaft (GA 683/9-1

When lithics hit bones: Evaluating the potential of a multifaceted experimental protocol to illuminate Middle Palaeolithic weapon technology

Recent zooarchaeological and isotope analyses have largely settled the debate surrounding Neanderthal hunting capacities, repeatedly demonstrating their successful acquisition of large ungulates. Nevertheless, the functional identification of individual tools as hunting weapons remains a methodological challenge. In-depth studies have focussed mainly on small subsets of lithic artefacts from selected assemblages assessing features of breakage patterns, retouch, shape and use wear. Studies focussing on associated hunting lesions are rarer and often focus on reconstructing very specific bone surface marks encountered in the archaeological record. This study aims to add to our understanding of the formation and characteristics of projectile impact marks (PIMs) on bone through a series of highly monitored, replicative experiments, using thrusting and throwing spears with replica Levallois points into two wild pig carcasses. In total, 152 shots were made, and for each a series of attributes was recorded, including velocity and location of impact. Subsequent quantitative analyses focussed on understanding the various factors underlying the formation of different types of projectile impact marks. These experiments demonstrate that PIM formation results from the properties of both the impacting projectile and bone element. PIMs can signal impacts caused by different delivery methods but only on some parts of the skeleton. These results are contextualised in relation to the occurrence and recognition of Palaeolithic PIMs and patterns of Neanderthal behaviour. These experiments are only a first step in improving the recognition of these signatures in the archaeological record and providing better insights into understanding of the mechanisms of Neanderthal hunting

Uneven Data Quality and the Earliest Occupation of Europe—the Case of Untermassfeld (Germany)

The database regarding the earliest occupation of Europe has increased significantly in quantity and quality of data points over the last two decades, mainly through the addition of new sites as a result of long-term systematic excavations and large-scale prospections of Early and early Middle Pleistocene exposures. The site distribution pattern suggests an ephemeral presence of hominins in the south of Europe from around 1 million years ago onward, with occasional short northward expansions along the western coastal areas when temperate conditions permitted. From around 600,000–700,000 years ago, Acheulean artefacts appear in Europe and somewhat later hominin presence seems to pick up, with more sites and now some also present in colder climatic settings. It is again only later, around 350,000 years ago, that the first sites show up in more continental, central parts of Europe, east of the Rhine. A series of recent papers on the Early Pleistocene palaeontological site of Untermassfeld (Thuringia, Germany) makes claims that are of great interest for studies of earliest Europe and are at odds with the described pattern: the papers suggest that Untermassfeld has yielded stone tools and humanly modified faunal remains, evidence for a 1 million years old hominin presence in European continental mid-latitudes, and additional evidence that hominins were well-established in Europe already around that time period. Here, we evaluate these claims and demonstrate that these studies are severely flawed in terms of data on provenance of the materials studied and in the interpretation of faunal remains and lithics as testifying to a hominin presence at the site. In actual fact, any reference to the Untermassfeld site as an archaeological one is unwarranted. Furthermore, it is not the only European Early Pleistocene site where inferred evidence for hominin presence is problematic. The strength of the spatiotemporal patterns of hominin presence and absence depends on the quality of the data points we work with, and database maintenance, including critical evaluation of new sites, is crucial to advance our knowledge of the expansions and contractions of hominin ranges during the Pleistocene.Human Origin

Mismatch between MIS 5e and the Last Interglacial (Eemian) of Central Europe.

Human Origin

Evaluating the incidence of hydrological processes during site formation through orientation analysis. A case study of the middle Palaeolithic Lakeland site of Neumark-Nord 2 (Germany)

Lacustrine localitieswere attractive environments for Palaeolithic hominins, since they provide a large and broad spectrumof resources. Moreover, they are excellent archives that allowfor high-resolution environmental, chronological and archaeological analyses. However, these deposits are often subject to complex formation and postdepositional factors, including water-related processes. Evaluating the influence of hydrological processes in site formation is thus essential to more accurately reconstruct the duration, intensity and types of hominin behaviour within these environments. In this paperwe present the orientation analysis of archaeological material from the Last Interglacial site Neumark-Nord 2, Germany. Orientation analysis was done using GIS to calculate the orientation of artefact from digital plans of the excavation surface, which were subsequently tested using circular statistics. The results of the orientation analysis are compared with a hydrological model to check the relation between preferred orientations and reconstructed areas of water flow and accumulation. Results suggest that low-energy hydrological processes could have affected certain areas of the find-bearing deposits at Neumark-Nord 2 but, overall, there is no evidence for either high-energy hydrological processes or a significant movement of parts of the archaeological assemblage.The project “Interglacial landscapes and Neanderthal adaptations in the Neumark-Nord region (Saxony-Anhalt)” is funded by the Deutsche Forchungsgemeinschaft (DFG) (GA 683/9-1)