Speciation of chilean Rhinocryptidae (Avian) based on their behaviour

The current classification of the chilean representatives of the passerine family Rhinocryptidae includes eight species. Three of them contain subspecies that don't exhibit clear differences. Moreover, differences among two lineages of _Scytalopus_ genera and two species of _Pteroptochos_ are cryptic. We propose a new methodology based on ecological and behavioural patterns in order to understand the concept of speciation in this group of birds. According to our results, we postulate that there is not a cut criteria to establish differences among three sister lineages of current classification. This way the methodology developed by us does not allow to establish divergence for a given common ancestor. Our methodology allows to establish comparison among previously determined phylogenetic lineages. Our results show how when integrating behaviour and ecological terms as biological traits next to morphological characters of the plumage, it allows us to conclude that there is decrease of the distances among sister lineages in the cluster tree

Detrital zircon SHRIMP U-Pb age study of the Cordillera Darwin Metamorphic Complex of Tierra del Fuego : sedimentary sources and implications for the evolution of the Pacific margin of Gondwana

The Cordillera Darwin Metamorphic Complex in the southernmost Andes includes a basement of probable Palaeozoic age, a mid-Jurassic and younger volcano-sedimentary cover, and a suite of Jurassic granites, all of which were jointly metamorphosed during the Cretaceous. Detrital zircon ages presented here show that some of the amphibolite-facies metamorphic rocks previously mapped as basement have a Jurassic protolith. Overall the detrital zircon age patterns for samples of the Cordillera Darwin basement differ from those of the Madre de Dios Terrane of the western Patagonian Andes with which they had been correlated; instead, they are more comparable with those from the Eastern Andes Metamorphic Complex, which apparently developed in a passive margin setting. The paucity of Cambrian detrital zircons indicates that the meta-igneous basement of the Magallanes foreland basin of central and northern Tierra del Fuego was not the main source of detritus for the protolith of the Cordillera Darwin Metamorphic Complex. The possibility is envisaged that the Magallanes Fagnano transform fault boundary between the Scotia and South America plates resulted from reactivation of an older, pre-Jurassic suture zone between the basement terranes of north–central Tierra del Fuego and Cordillera Darwin

Subduction Duration and Slab Dip

The dip angles of slabs are among the clearest characteristics of subduction zones, but the factors that control them remain obscure. Here, slab dip angles and subduction parameters, including subduction duration, the nature of the overriding plate, slab age, and convergence rate, are determined for 153 transects along subduction zones for the present day. We present a comprehensive tabulation of subduction duration based on isotopic ages of arc initiation and stratigraphic, structural, plate tectonic and seismic indicators of subduction initiation. We present two ages for subduction zones, a long‐term age and a reinitiation age. Using cross correlation and multivariate regression, we find that (1) subduction duration is the primary parameter controlling slab dips with slabs tending to have shallower dips at subduction zones that have been in existence longer; (2) the long‐term age of subduction duration better explains variation of shallow dip than reinitiation age; (3) overriding plate nature could influence shallow dip angle, where slabs below continents tend to have shallower dips; (4) slab age contributes to slab dip, with younger slabs having steeper shallow dips; and (5) the relations between slab dip and subduction parameters are depth dependent, where the ability of subduction duration and overriding plate nature to explain observed variation decreases with depth. The analysis emphasizes the importance of subduction history and the long‐term regional state of a subduction zone in determining slab dip and is consistent with mechanical models of subduction

Paleogene building of the Bolivian Orocline: Tectonic restoration of the central Andes in 2-D map view

International audienceUsing available information on the magnitude and age of tectonic shortening, as well as paleomagnetically determined tectonic rotations, we have run a series of 2-D map view restorations of the central Andes. Neogene shortening in the foreland belt induced only slight orogenic curvature of the central Andes. The constraints on the ages of the large observed fore-arc rotations (average of 37° counterclockwise in southern Peru and 29° clockwise in northern Chile) indicate that the Bolivian Orocline formed during the Eocene-Oligocene as a consequence of differential shortening focused in the Eastern Cordillera. To minimize local block rotations along the fore arc, the restoration that best fits the central Andean rotation pattern requires about 400 km of total (Paleogene plus Neogene) shortening near the Arica bend. This value corresponds to the upper bound of estimates of maximum horizontal shortening for the central Andes. Along-strike variations in horizontal shortening in the back arc induced bending of the continental margin, block rotations with fore-arc along-strike extension, and/or orogen-parallel transport of upper crustal material toward the symmetry axis of the orocline

On Entangled Multi-particle Systems in Bohmian Theory

Arguments are presented to show that in the case of entangled systems there are certain difficulties in implementing the usual Bohmian interpretation of the wave function in a straightforward manner. Specific examples are given.Comment: 7 page

Paleomagnetism of Permian and Triassic Rocks, Central Chilean Andes

The first paleomagnetic data from Permian and Triassic formations west of the Andean divide are presented. Four formations of Permian or Triassic age in the central Chilean Andes have been investigated: two are located in the coastal ranges, and two are in the main cordillera. Of the formations in the main cordillera (Pastos Blancos and Matahuaico formations), only the Pastos Blancos Formation has yielded characteristic directions. While a fold test is absent, magnetizations are most likely secondary and yield pre-tilt corrected concordant inclinations, but yield declinations discordant 30° clockwise in comparison to the South American apparent polar wander path. Both formations from the coastal ranges (Cifuncho and Pichidangui formations) yielded stable directions. Postfolding magnetizations in the Cifuncho Formation also show declinations discordant 30° clockwise and concordant inclinations. The Pichidangui Formation has two stable components: one of postfolding age is concordant to apparent polar wander path data, and one of probable prefolding (Late Triassic) age is concordant in declination, but discordant in inclination. Further work is needed to better define the prefolding magnetizations in the Pichidangui Formation, but at present these preliminary results are the first paleomagnetic signs of displaced terranes along the Pacific margin of Chile. If correct, the results suggest that the Pichidangui Formation was some 15° of latitude farther south during the Late Triassic and had likely moved northward to its present latitudinal position with respect to cratonic South America by Middle to Late Jurassic

Jurassic tectonics in Argentina and Chile: Extension, oblique subduction, rifting, drift and collisions?

The Jurassic history of southern South America shows a complex geologic evolution which is the result of different processes that began along the western Gondwana margin during the initial stages of Pangea breakup. Andean subduction along the Pacific continental margin began in the Early Jurassic, after a period of continental-scale extension and rifting, which peaked by the end of the Triassic in central and northern Argentina and Chile. Renewal of subduction was the result of an episode of ocean growth along a series of spreading centers between North and South America when the separation of these continents began as a consequence of the activity of the Central Atlantic Magmatic Province hotspot. Motion along these spreading centers produced a component of oblique, SE-directed subduction along the western margin of South America and the reactivation of inherited orthogonal structural features as the N70°E trending Huincul ridge in the Neuquén Basin that was uplifted during Jurassic times. Subduction along the north-south trending Argentine-Chilean continental margin acelerated during the break-up between West and East Gondwana soon after the opening of the Indian Ocean, linked to the Karoo hot-spot. Subduction took place under extensional conditions probably associated with a negative trench roll-back, leading to the formation of a magmatic arc along the Coast Ranges from southern Peru to central Chile and, to the east, the Arequipa, Tarapacá and Neuquén extensional back-arc basins. In northern Patagonia, early Jurassic arc related magmatism occurred to the east of the present day Andean Cordillera along the short-lived (190-170 Ma) Subcordilleran Batholith and the associated Liassic intra arc basin. Arc magmatism ceased in northern Patagonia at ca 170 Ma to be replaced by huge volumes of Early to Middle Jurassic rhyolites and dacites of the Chon-Aike Large Igneous province produced as a result of crustal melting in an overheated crust during the initial stages of Gondwana breakup. Early rifting during Middle-Late Jurassic times took place in the Cañadón Asfalto Basin and the Late Jurassic Río Guenguel, Río Mayo and Río Senguerr basins, orthogonal to the continental margin as a consequence of the Weddell Sea opening. Acid magmatism was associated with widespread extension and culminated in the opening of the ocean-floored Rocas Verdes Basin. The causes of the cessation of magmatism in the Subcordilleran Batholith, the origin of the Chon Aike LIP and the rotation of the magmatic front towards the Patagonian Batholith around 150 Ma are still not well understood. Hypothesis linking this mutating tectonic scenario to the collision of exotic terranes against the Pacific margin of Patagonia during the early to middle Jurassic should be taken into consideration.Fil:Ramos, V.A. Universidad de Buenos Aires. Facultad de Ciencias Exactas y Naturales; Argentina

Deterioration of the Gαo Vomeronasal Pathway in Sexually Dimorphic Mammals

In mammals, social and sexual behaviours are largely mediated by the vomeronasal system (VNS). The accessory olfactory bulb (AOB) is the first synaptic locus of the VNS and ranges from very large in Caviomorph rodents, small in carnivores and ungulates, to its complete absence in apes, elephants, most bats and aquatic species. Two pathways have been described in the VNS of mammals. In mice, vomeronasal neurons expressing Gαi2 protein project to the rostral portion of the AOB and respond mostly to small volatile molecules, whereas neurons expressing Gαo project to the caudal AOB and respond mostly to large non-volatile molecules. However, the Gαo-expressing pathway is absent in several species (horses, dogs, musk shrews, goats and marmosets) but no hypotheses have been proposed to date to explain the loss of that pathway. We noted that the species that lost the Gαo pathway belong to Laurasiatheria and Primates lineages, both clades with ubiquitous sexual dimorphisms across species. To assess whether similar events of Gαo pathway loss could have occurred convergently in dimorphic species we studied G-protein expression in the AOB of two species that independently evolved sexually dimorphic traits: the California ground squirrel Spermophilus beecheyi (Rodentia; Sciurognathi) and the cape hyrax Procavia capensis (Afrotheria; Hyracoidea). We found that both species show uniform expression of Gαi2-protein throughout AOB glomeruli, while Gαo expression is restricted to main olfactory glomeruli only. Our results suggest that the degeneration of the Gαo-expressing vomeronasal pathway has occurred independently at least four times in Eutheria, possibly related to the emergence of sexual dimorphisms and the ability of detecting the gender of conspecifics at distance