Temperate Subantarctic Forests: A Huge Natural Laboratory

In the southern tip of South America, temperate subantarctic forests develop on both sides of the Andes mountain range. Known in Argentina as Andean-Patagonian forests, these ecosystems cover a narrow but long latitudinal strip, from 35° S (at Maule River, in Chile) to 55° S (in the southern extreme of the Tierra del Fuego archipelago, in Chile and Argentina). Their current geographic discontinuity from the other forests of South America is a remarkable feature, for which they have been considered a biogeographic island (Armesto et al. 1995). The sclerophyllous scrub and the high-Andean-steppe border the Subantarctic forests to the north; the Patagonian steppe, in Argentina, confines its development to the east, and the ocean marks its western and southern edges. This geographic isolation dates back to the Oligocene (about 23–33 My ago), when South America began to drift northward; before that, the continents of the southern hemisphere were connected as Gondwanaland (Markgraf et al. 1996). Since its origin in the late Cretaceous (ca. 90 My ago; Dettmann et al. 1990) till the separation of the continents, floristic interchange endured. This explains the relationships with other southern forests like those of Tasmania and New Zealand (i.e., disjunct distribution of genus like Araucaria, Aristotelia, Blechnum, Discaria, Lomatia, Nothofagus, Podocarpus, among others; Veblen et al. 1996). In addition, abundant neotropical elements characteristics of the “Yungas” in NE Argentina (e.g., Azara, Chusquea, Crinodendron, Drimys, Escallonia among others) reflect a history of recurrent connections with ecosystems of lower latitudes (Arroyo et al. 1995).Estación Experimental Agropecuaria BarilocheFil: Marchelli, Paula. Instituto Nacional de Tecnología Agropecuaria (INTA). Estación Experimental Agropecuaria Bariloche; ArgentinaFil: Marchelli, Paula. Consejo Nacional de Investigaciones Cientificas y Tecnicas. Instituto de Investigaciones Forestales y Agropecuarias Bariloche; ArgentinaFil: Pastorino, Mario Juan. Instituto Nacional de Tecnología Agropecuaria (INTA). Estación Experimental Agropecuaria Bariloche; ArgentinaFil: Pastorino, Mario Juan. Consejo Nacional de Investigaciones Cientificas y Tecnicas. Instituto de Investigaciones Forestales y Agropecuarias Bariloche; ArgentinaFil: Gallo, Leonardo Ariel. Instituto Nacional de Tecnología Agropecuaria (INTA). Estación Experimental Agropecuaria Bariloche; ArgentinaFil: Gallo, Leonardo Ariel. Consejo Nacional de Investigaciones Cientificas y Tecnicas. Instituto de Investigaciones Forestales y Agropecuarias Bariloche; Argentin