An Ozone-Modified Refractive Index for Vertically Propagating Planetary Waves

[1] An ozone-modified refractive index (OMRI) is derived for vertically propagating planetary waves using a mechanistic model that couples quasigeostrophic potential vorticity and ozone volume mixing ratio. The OMRI clarifies how wave-induced heating due to ozone photochemistry, ozone transport, and Newtonian cooling (NC) combine to affect wave propagation, attenuation, and drag on the zonal mean flow. In the photochemically controlled upper stratosphere, the wave-induced ozone heating (OH) always augments the NC, whereas in the dynamically controlled lower stratosphere, the wave-induced OH may augment or reduce the NC depending on the detailed nature of the wave vertical structure and zonal mean ozone gradients. For a basic state representative of Northern Hemisphere winter, the wave-induced OH can increase the planetary wave drag by more than a factor of two in the photochemically controlled upper stratosphere and decrease it by as much as 25% in the dynamically controlled lower stratosphere. Because the zonal mean ozone distribution appears explicitly in the OMRI, the OMRI can be used as a tool for understanding how changes in stratospheric ozone due to solar variability and chemical depletion affect stratosphere-troposphere communication

A New Pathway for Communicating the 11-year Solar Cycle Signal to the QBO

[1] The response of the equatorial quasi-biennial oscillation (QBO) to zonal-mean ozone perturbations consistent with the 11-year solar cycle is examined using a 2 1/2 dimensional model of the tropical stratosphere. Unique to this model are wave-ozone feedbacks, which provide a new, nonlinear pathway for communicating solar variability effects to the QBO. Model simulations show that for zonal-mean ozone perturbations representative of solar maximum (minimum), the diabatic heating due to the wave-ozone feedbacks is primarily responsible for driving a slightly stronger (weaker) QBO circulation and producing a slightly shorter (longer) QBO period. These results, which are explained via an analytical analysis of the divergence of Eliassen-palm flux, are in general agreement with observations of quasi-decadal variability of the QBO

An Examination of Anomalously Low Column Ozone in the Southern Hemisphere Midlatitudes During 1997

[1] Observations from both ground-based and satellite instruments show record low column ozone abundance between 20°S and 40°S during 1997. The 1997 monthly averaged column ozone from the Total Ozone Mapping Spectrometer (TOMS) is up to 25 Dobson units (DU) lower than the TOMS climatological mean (1979–1996) and up to 20 DU below the previous record low values. Observations from the Halogen Occultation Experiment show that below average ozone concentrations during 1997 were confined primarily to the lower stratosphere. Residual circulation statistics calculated from the United Kingdom Meteorological Office temperature analyses indicate that circulation anomalies during 1997 can account for ∼5–10 DU/month decrease in column ozone between 20°S and 50°S. At these latitudes during 1997, structural characteristics of the ozone and residual circulation fields both suggest a connection with the equatorial quasi-biennial oscillation

The Attentional Routing Circuit: Receptive Field Modulation Through Nonlinear Dendritic Interactions

We present a model of attentional routing called the Attentional Routing Circuit (ARC) that extends an existing model of spiking neurons with dendritic nonlinearities. Specifically, we employ the Poirazi et al. (2003) pyramidal neuron in a population coding framework. ARC demonstrates that the dendritic nonlinearities can be exploited to result in selective routing, with a decrease in the number of cells needed by a factor of ~5 as compared with a linear dendrite model.

Routing of attended information occurs through the modulation of feedforward visual signals by a cortical control signal specifying the location and size of the attended target. The model is fully specified in spiking single cells. Our approach differs from past work on shifter circuits by having more efficient control, and using a more biologically detailed substrate. Our approach differs from existing models that use gain fields by providing precise hypotheses about how the control signals are generated and distributed in a hierarchical model in spiking neurons. Further, the model accounts for numerous experimental findings regarding the timing, strength and extent of attentional modulation in ventral stream areas, and the perceived contrast enhancement of attended stimuli.

To further demonstrate the plausibility of ARC, it is applied to the attention experiments of Womelsdorf et al. (2008) and tested in detail. For the simulations, the model has only two free parameters that influence its ability to match the experimental data, and without fitting, we show that it can account for the experimental observations of changes in receptive field (RF) gain and position with attention in macaques. In sum, the model provides an explanation of RF modulation as well as testable predictions about nonlinear cortical dendrites and attentional changes of receptive field properties

Long slit spectroscopy of NH2 in comets Halley, Wilson, and Nishikawa-Takamizawa-Tago

Long-slit spectra of comets Halley, Wilson and Nishikawa-Takamizawa-Tago were obtained with the 3.9 meter Anglo-Australian Telescope. Spectra of comets Halley and Wilson were obtained with the IPCS at a spectral resolution of 0.5 A and a spatial resolution of 10(exp 3) km. Spectra of comets Wilson and Nishikawa-Takamizawa-Tago were obtained with a CCD at a spectral resolution of 1.5 A and a spatial resolution of approximately 3 x 10(exp 3) km. Surface brightness profiles for NH2 were extracted from the long-slit spectra of each comet. The observed surface brightness profiles extend along the slit to approximately 6 x 10(exp 4) km from the nucleus in both sunward and tailward directions. By comparing surface distribution calculated from an appropriate coma model with observed surface brightness distributions, the photodissociation timescale of the parent molecule of NH2 can be inferred. The observed NH2 surface brightness profiles in all three comets compares well with a surface brightness profile calculated using the vectorial model, an NH3 photodissociation timescale of 7 x 10(exp 3) seconds, and an NH2 photodissociation timescale of 34,000 seconds

CO Line Emission and Absorption from the HL Tau Disk: Where is all the dust?

We present high-resolution infrared spectra of HL Tau, a heavily embedded young star. The spectra exhibit broad emission lines of hot CO gas as well as narrow absorption lines of cold CO gas. The column density for this cooler material (7.5+/-0.2 x 10^18 cm-2) indicates a large column of absorbing gas along the line of sight. In dense interstellar clouds, this column density of CO gas is associated with Av~52 magnitudes. However, the extinction toward this source (Av~23) suggests that there is less dust along the line of sight than inferred from the CO absorption data. We discuss three possibilities for the apparent paucity of dust along the line of sight through the flared disk: 1) the dust extinction has been underestimated due to differences in circumstellar grain properties, such as grain agglomeration; 2) the effect of scattering has been underestimated and the actual extinction is much higher; or (3) the line of sight through the disk is probing a gas-rich, dust-depleted region, possibly due to the stratification of gas and dust in a pre-planetary disk.Comment: To be published in The Astrophysical Journa

Effects of Planetary Wave-breaking on the Seasonal Variation of Total Column Ozone

The effects of planetary wave breaking on the seasonal variation of total column ozone are investigated using a zonally averaged chemical-radiative-transport model of the atmosphere. The planetary wave breaking effects of zonal wavenumbers k=1 and k=2 are significant in the middle latitude stratosphere during Northern Hemisphere (NH) winter, whereas only wave k=1 is important during Southern Hemisphere (SH) winter. The mixing and induced meridional circulation due to the planetary wave breaking increases the seasonal variation of total column ozone in NH (SH) middle latitudes by ∼20% (∼10%)

On the predominant nonlinear response of the extratropical atmosphere to meridional shifts of the Gulf Stream

Author Posting. © American Meteorological Society, 2017. This article is posted here by permission of American Meteorological Society for personal use, not for redistribution. The definitive version was published in Journal of Climate 30 (2017): 9679-9702, doi:10.1175/JCLI-D-16-0707.1.The North Atlantic atmospheric circulation response to the meridional shifts of the Gulf Stream (GS) path is examined using a large ensemble of high-resolution hemispheric-scale Weather Research and Forecasting Model simulations. The model is forced with a broad range of wintertime sea surface temperature (SST) anomalies derived from a lag regression on a GS index. The primary result of the model experiments, supported in part by an independent analysis of a reanalysis dataset, is that the large-scale quasi-steady North Atlantic circulation response is remarkably nonlinear about the sign and amplitude of the SST anomaly chosen over a wide range of GS shift scenarios. The nonlinear response prevails over the weak linear response and resembles the negative North Atlantic Oscillation (NAO), the leading intrinsic mode of variability in the model and the observations. Further analysis of the associated dynamics reveals that the nonlinear responses are accompanied by the shift of the North Atlantic eddy-driven jet, which is reinforced, with nearly equal importance, by the high-frequency transient eddy feedback and the low-frequency wave-breaking events. Additional sensitivity simulations confirm that the nonlinearity of the circulation response is a robust feature found over the broad parameter space encompassing not only the varied SST but also the absence/presence of tropical influence, the varying lateral boundary conditions, and the initialization scheme. The result highlights the fundamental importance of the intrinsically nonlinear transient eddy dynamics and the eddy–mean flow interactions in generating the nonlinear downstream response to the meridional shifts in the Gulf Stream.The authors are grateful for the support from NASA (NNX13AM59G) and the NSF (AGS-1355339, OCE-1419235).2018-05-0

Mitofusins and OPA1 Mediate Sequential Steps in Mitochondrial Membrane Fusion

Mitochondrial fusion requires the coordinated fusion of the outer and inner membranes. Three large GTPases—OPA1 and the mitofusins Mfn1 and Mfn2—are essential for the fusion of mammalian mitochondria. OPA1 is mutated in dominant optic atrophy, a neurodegenerative disease of the optic nerve. In yeast, the OPA1 ortholog Mgm1 is required for inner membrane fusion in vitro; nevertheless, yeast lacking Mgm1 show neither outer nor inner membrane fusion in vivo, because of the tight coupling between these two processes. We find that outer membrane fusion can be readily visualized in OPA1-null mouse cells in vivo, but these events do not progress to inner membrane fusion. Similar defects are found in cells lacking prohibitins, which are required for proper OPA1 processing. In contrast, double Mfn-null cells show neither outer nor inner membrane fusion. Mitochondria in OPA1-null cells often contain multiple matrix compartments bounded together by a single outer membrane, consistent with uncoupling of outer versus inner membrane fusion. In addition, unlike mitofusins and yeast Mgm1, OPA1 is not required on adjacent mitochondria to mediate membrane fusion. These results indicate that mammalian mitofusins and OPA1 mediate distinct sequential fusion steps that are readily uncoupled, in contrast to the situation in yeast